Youth and education in Eugene, OR; UC Berkeley graduate study in botany, 1930-1934; teaching at Washington State College, 1934-1937; professor of botany, UC Berkeley, 1937-1976; William Setchell and Willis Jepson, the Bay Area Biosystematists; work as a plant taxonomist: seminal papers, colleagues Mildred Mathias, Marion Cave, Reed Rollins, prominent graduate students, worldwide research on Umbelliferae; Harvard, 1947-1948; sabbatical in Chile and Peru, 1953-1954; Academic Senate service, including Committee on Budget and Interdepartmental Relations; faculty role in university governance; vice chancellor, 1962-1965, under Edward Strong and Martin Meyerson: Free Speech Movement, relations with President Clark Kerr, student attitudes, faculty responses; director, UC Herbarium, 1963-1975; president, California Academy of Science, 1975-1978; emeritus research.